Flehmen Grimace (Panthera leo), Katavi National Park, Tanzania

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Description

A sub-adult male makes the classic facial gesture known as the Flehmen Grimace as he tests the urine from a pride female for signs of oestrus using the vomeronasal organ. Such behaviour will encourage the pride male to drive him out of the pride for years of wandering as a nomad before he is big and strong enough to hold a pride of his own

Photographer: Nick Greaves
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